The extreme, workerless inquilines.

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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Sonntag 17. Oktober 2021, 12:06

I was wandering yesterday and today in AntWiki and AntWeb and made some (late) observations about the hosts of some Pheidole species. So I did put them in my list and made some comments about them. Also I made remarks about the changes of genus as ", by" and then the name and date from the change. Hope you enjoy it!

P.s. the reference Casadei-Ferreira et al., 2020 refers to:

Casadei-Ferreira, A.; Economo, E. P.; Feitosa, R. M. 2020. Additions to the taxonomy of Pheidole (Hymenoptera: Formicidae) from the southern grasslands of Brazil. Revista Brasileira de Entomologia 64 (4):21 pp.

http://antcat.org/documents/8034/casade ... eidole.pdf
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Sonntag 17. Oktober 2021, 12:10

The extreme, workerless inquilines.

Teleutomyrmex Kutter, 1950

01) Teleutomyrmex schneideri Kutter, 1950
(= Tetramorium schneideri (Kutter, 1950), by Ward, Brady, Fisher & Schultz, 2015 ("2014"))
(not Tetramorium schneideri Emery, 1898)
(= Tetramorium inquilinum Ward, Brady, Fisher & Schultz, 2015 ("2014"))
02) Teleutomyrmex kutteri Tinaut, 1990
(= Tetramorium kutteri (Tinaut, 1990), by Ward, Brady, Fisher & Schultz, 2015 ("2014"))
(not Tetramorium semilaeve kutteri Santschi, 1927)
03) Teleutomyrmex seiferti Kiran & Karaman, in Kiran, et al. 2017
(= Tetramorium seiferti (Kiran & Karaman, in Kiran, et al. 2017), by analogy)
04) Teleutomyrmex buschingeri Lapeva-Gjonova, in Kiran, et al. 2017
(= Tetramorium buschingeri (Lapeva-Gjonova, in Kiran, et al. 2017), by analogy)

Not yet described species of extreme, workerless inquiline, from the genus Teleutomyrmex Kutter, 1950

05) The new, undescribed species from Teleutomyrmex Kutter, 1950 from Farab, Turkmenistan… See Dlussky, et al. 1990.

Anergates Forel, 1874

06) Anergates atratulus (Schenck, 1852), by Forel, 1874
(= Myrmica atratula Schenck, 1852)
[Also described as new by Schenck, 1853b]
(= Tetramorium atratulum (Schenck, 1852), by Mayr, 1855)
[= Tomognathus atratulus (Schenck, 1852), by Mayr, 1863 following Mayr, 1861, obsolete combination.]
(= Tetramorium atratulum (Schenck, 1852), by Ward, Brady, Fisher & Schultz, 2015 ("2014"))
07) Anergates friedlandi Creighton, 1934
[= Tetramorium friedlandi (Creighton, 1934), by analogy]

Tetramorium Mayr, 1855
(Only a few species in a big genus.)

08) Tetramorium microgyna Santschi, 1918
09) Tetramorium parasiticum Bolton, 1980

Pheidole Westwood, 1839
(Only a few species in a big genus.)

10) Pheidole neokohli Wilson, 1984
(= Pheidole kohli (Wasmann, 1915), by Wilson, 1984)
(= Anergatides kohli Wasmann, 1915)
(not Pheidole kohli Mayr, 1901)
11) Pheidole acutidens (Santschi, 1922), by Wilson, 1984
(= Bruchomyrma acutidens Santschi, 1922)
12) Pheidole argentina (Bruch, 1932), by Wilson, 1984
(= Gallardomyrma argentina Bruch, 1932)
13) Pheidole parasitica Wilson, 1984

Excluded from the extreme, workerless inquilines
Once this species was included in the extreme, workerless inquilines but now it is considered to be a workerless inquiline without extreme reductions, e.g. no pupoid males but normal ones. The decision to exclude it was made by Edward Osborne Wilson in 1984 in a study of the inquilines in the genus Pheidole Westwood, 1839.

Pheidole Westwood, 1839
(Only one species in a big genus.)

14) Pheidole kusnezovi Wilson, 2003
(= Pheidole symbiotica (Kusnezov, 1952), by Wilson, 1984)
(= Eriopheidole symbiotica Kusnezov, 1952)
(not Pheidole symbiotica Wasmann, 1909)

Distribution

01) Europe (Alps, Pyrenees and Northern Spain)
02) Europe (Southern Iberia)
03) Turkey (Anatolia)
04) Europe (Southern Balkans)

05) Turkmenistan

06) Europe
07) North America

08) Southern Africa
09) Southern Africa

10) Central Africa
11) South America
12) South America
13) India

14) South America

Host species

01), 02), 03), 04), 05), 06), 07), 08) and 09) Certain species of the genus Tetramorium Mayr, 1855

01) T. alpestre Steiner, Schlick-Steiner & Seifert, 2010 and T. impurum (Förster, 1850)
and maybe T. caespitum (Linnaeus, 1758)?
02) T. cf. caespitum (Linnaeus, 1758)
03) T. cf. chefketi Forel, 1911
04) T. cf. chefketi Forel, 1911

05) A species from the genus Tetramorium Mayr, 1855…

06) T. impurum (Förster, 1850), T. caespitum (Linnaeus, 1758), T. immigrans Santschi, 1927
and T. moravicum Kratochvil, 1941, T. diomedeum Emery, 1908, T. chefketi Forel, 1911
07) T. immigrans Santschi, 1927

08) T. sericeiventre Emery, 1877 and T. sepositum Santschi, 1918
09) T. avium Bolton, 1980

10), 11), 12), 13) and 14) Certain species of the genus Pheidole Westwood, 1839

10) P. megacephala (Fabricius, 1793) subsp. melancholica Santschi, 1912
11) P. strobeli Emery, 1906
12) P. nitidula Emery, 1888
13) P. indica Mayr, 1879

14) P. obscurior Forel, 1886

A remark about synonymy

Anergates friedlandi Creighton, 1934 is now a synonym from Anergates atratulus (Schenck, 1852), more precisely an introduced form in North Americe. So, the name is Anergates atratulus (Schenck, 1852) or, if you follow Ward et al. 2015 ("2014"), Tetramorium atratulum (Schenck, 1852)...

Synonyms of host species

- T. caespitum (Linnaeus, 1758) (= Formica caespitum Linnaeus, 1758)
- T. impurum (Förster, 1850) (= Myrmica impura Förster, 1850)
- P. megacephala (Fabricius, 1793) (= Formica megecephala Fabricius, 1793)
- P. megacephala (Fabricius, 1793) subsp. melancholica Santschi, 1912 was originally described as P. punctulata Mayr, 1866 st. melancholica Santschi, 1912
- The host species P. strobeli Emery, 1906 was, in 1922, originally described as a subspecies of an other species, nl. P. nitidula Emery, 1888 subsp. richteri Forel, 1909 (= the protonym P. perversa Forel, 1908 subsp. richteri Forel, 1909) and known as P. strobeli Emery, 1906 subsp. richteri Forel, 1909 since Santschi, 1916. The species P. strobeli Emery, 1906 was revived from synonymy (with P. nitidula Emery, 1888) and the subspecies was synonymized with it in Casadei-Ferreira et al., 2020.
- P. obscurior Forel, 1886 was revived from synonymy with the 1 page older P. susannae Forel, 1886 and this in Casadei-Ferreira et al., 2020.
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Sonntag 17. Oktober 2021, 17:26

The two pdf-files:

The extreme, workerless inquilines 44.pdf
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Official literature lists 41.pdf
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Mittwoch 20. Oktober 2021, 18:27

- The host species P. strobeli Emery, 1906 was, in 1922, originally described as a subspecies of an other species, nl. P. nitidula Emery, 1888 subsp. richteri Forel, 1909 (= the protonym P. perversa Forel, 1908 subsp. richteri Forel, 1909) and known as P. strobeli Emery, 1906 subsp. richteri Forel, 1909 since Santschi, 1916. The species P. strobeli Emery, 1906 was revived from synonymy (with P. nitidula Emery, 1888) and the subspecies was synonymized with it in Casadei-Ferreira et al., 2020.

This is changed into a better version:

- The host species P. strobeli Emery, 1906 was, in 1922, described as a subspecies of an other species, nl. P. nitidula Emery, 1888 subsp. richteri Forel, 1909 (a much used junior synonym of the older P. perversa Forel, 1908 subsp. richteri Forel, 1909) and known as P. strobeli Emery, 1906 subsp. richteri Forel, 1909 since Santschi, 1916. The species P. strobeli Emery, 1906 was revived from synonymy (with P. nitidula Emery, 1888 since Wilson, 2003) and the subspecies was synonymized with it in Casadei-Ferreira et al., 2020.
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Sonntag 31. Oktober 2021, 13:11

The extreme, workerless inquilines 45.pdf
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Samstag 6. November 2021, 14:43

I made a few changes in the lists.

First in "The extreme, workerless inquilines":

"T. moravicum Kratochvil, 1941"
I did change this. "Kratochvil" or "Kratochvil, in Novák & Sadil, 1941" you don't see much, most it is "Novák & Sadil". So I made this:
"T. moravicum [Kratochvil, in] Novák & Sadil, 1941"

And I made "- The host species P. strobeli Emery, 1906 was, in 1922, described as a subspecies of an other species, nl. P. nitidula Emery, 1888 subsp. richteri Forel, 1909 (a much used junior synonym of the older P. perversa Forel, 1908 subsp. richteri Forel, 1909) and known as P. strobeli Emery, 1906 subsp. richteri Forel, 1909 since Santschi, 1916. The species P. strobeli Emery, 1906 was revived from synonymy (with P. nitidula Emery, 1888 since Wilson, 2003) and the subspecies was synonymized with it in Casadei-Ferreira et al., 2020." more understandible, nl. this:
"- The host species P. strobeli Emery, 1906 was known, during much of its history, as a subspecies of an other species, nl. P. nitidula Emery, 1888 subsp. richteri Forel, 1909 (a much used junior synonym of the older P. perversa Forel, 1908 subsp. richteri Forel, 1909) and known, in 1922, for a while as P. strobeli Emery, 1906 subsp. richteri Forel, 1909, this since Santschi, 1916. The species P. strobeli Emery, 1906 was revived from synonymy (with P. nitidula Emery, 1888 since Wilson, 2003) and the subspecies was synonymized with it in Casadei-Ferreira, Economo, Feitosa, 2020."

And an addition to the references of Anergates, but separate from them:
Schlick-Steiner, B. C., Steiner, F. M., Sanetra, M., Heller, G., Stauffer, C., Christian, E., Seifert, B., 2005, “Queen size dimorphism in the ant Tetramorium moravicum (Hymenoptera, Formicidae): morphometric, molecular genetic and experimental evidence.” Insectes Sociaux, vol. 52, p. 186-193.
Schulz, A., 1996, “Tetramorium rhenanum nov. spec. vom "Mittleren Rheintal" in Deutschland (Hymenoptera: Formicidae).” Linzer Biologische Beiträge, vol. 28, p. 391-412.

And a few little things, more uniform....
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Samstag 6. November 2021, 14:45

Official literature lists 49.pdf
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The extreme, workerless inquilines 49.pdf
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Sonntag 28. November 2021, 20:21

The extreme, workerless inquilines.

Teleutomyrmex Kutter, 1950

01) Teleutomyrmex schneideri Kutter, 1950
(= Tetramorium schneideri (Kutter, 1950), by Ward, Brady, Fisher, Schultz, 2015 ("2014"))
(not Tetramorium schneideri Emery, 1898)
(= Tetramorium inquilinum Ward, Brady, Fisher, Schultz, 2015 ("2014"))
02) Teleutomyrmex kutteri Tinaut, 1990
(= Tetramorium kutteri (Tinaut, 1990), by Ward, Brady, Fisher, Schultz, 2015 ("2014"))
(not Tetramorium semilaeve kutteri Santschi, 1927)
03) Teleutomyrmex seiferti Kiran & Karaman, in Kiran, Karaman, Lapeva-Gjonova, Aksoy, 2017
(= Tetramorium seiferti (Kiran & Karaman, in Kiran, Karaman, Lapeva-Gjonova, Aksoy, 2017), by analogy)
04) Teleutomyrmex buschingeri Lapeva-Gjonova, in Kiran, Karaman, Lapeva-Gjonova, Aksoy, 2017
(= Tetramorium buschingeri (Lapeva-Gjonova, in Kiran, Karaman, Lapeva-Gjonova, Aksoy, 2017), by analogy)

Not yet described species of extreme, workerless inquiline, from the genus Teleutomyrmex Kutter, 1950

05) The new, undescribed species from Teleutomyrmex Kutter, 1950 from Farab, Turkmenistan… See Dlussky, Soyunov, Zabelin, 1990 [“1989”].

Anergates Forel, 1874

06) Anergates atratulus (Schenck, 1852), by Forel, 1874
(= Myrmica atratula Schenck, 1852)
[Also described as new by Schenck, 1853b]
(= Tetramorium atratulum (Schenck, 1852), by Mayr, 1855)
[= Tomognathus atratulus (Schenck, 1852), by Mayr, 1863 following Mayr, 1861, obsolete combination.]
(= Tetramorium atratulum (Schenck, 1852), by Ward, Brady, Fisher, Schultz, 2015 ("2014"))
07) Anergates friedlandi Creighton, 1934
[= Tetramorium friedlandi (Creighton, 1934), by analogy]

Tetramorium Mayr, 1855
(Only a few species in a big genus.)

08) Tetramorium microgyna Santschi, 1918
09) Tetramorium parasiticum Bolton, 1980

Pheidole Westwood, 1839
(Only a few species in a big genus.)

10) Pheidole neokohli Wilson, 1984
(= Pheidole kohli (Wasmann, 1915), by Wilson, 1984)
(= Anergatides kohli Wasmann, 1915)
(not Pheidole kohli Mayr, 1901)
11) Pheidole acutidens (Santschi, 1922), by Wilson, 1984
(= Bruchomyrma acutidens Santschi, 1922)
12) Pheidole argentina (Bruch, 1932), by Wilson, 1984
(= Gallardomyrma argentina Bruch, 1932)
13) Pheidole parasitica Wilson, 1984

Excluded from the extreme, workerless inquilines
Once this species was included in the extreme, workerless inquilines but now it is considered to be a workerless inquiline without extreme reductions, e.g. no pupoid males but normal ones. The decision to exclude it was made by Edward Osborne Wilson in 1984 in a study of the inquilines in the genus Pheidole Westwood, 1839.

Pheidole Westwood, 1839
(Only one species in a big genus.)

14) Pheidole kusnezovi Wilson, 2003
(= Pheidole symbiotica (Kusnezov, 1952), by Wilson, 1984)
(= Eriopheidole symbiotica Kusnezov, 1952)
(not Pheidole symbiotica Wasmann, 1909)

Distribution

01) Europe (Alps, Pyrenees and Northern Spain)
02) Europe (Southern Iberia)
03) Turkey (Anatolia)
04) Europe (Southern Balkans)

05) Turkmenistan

06) Europe
07) North America

08) Southern Africa
09) Southern Africa

10) Central Africa
11) South America
12) South America
13) India

14) South America

Host species

01), 02), 03), 04), 05), 06), 07), 08) and 09) Certain species of the genus Tetramorium Mayr, 1855

01) T. alpestre Steiner, Schlick-Steiner & Seifert, 2010 and T. impurum (Förster, 1850)
and maybe T. caespitum (Linnaeus, 1758)?
02) T. cf. caespitum (Linnaeus, 1758)
03) T. cf. chefketi Forel, 1911
04) T. cf. chefketi Forel, 1911

05) A species from the genus Tetramorium Mayr, 1855…

06) T. impurum (Förster, 1850), T. caespitum (Linnaeus, 1758), T. immigrans Santschi, 1927
and T. moravicum [Kratochvil, in] Novák & Sadil, 1941, T. diomedeum Emery, 1908, T. chefketi Forel, 1911
07) T. immigrans Santschi, 1927

08) T. sericeiventre Emery, 1877 and T. sepositum Santschi, 1918
09) T. avium Bolton, 1980

10), 11), 12), 13) and 14) Certain species of the genus Pheidole Westwood, 1839

10) P. megacephala (Fabricius, 1793) subsp. melancholica Santschi, 1912
11) P. strobeli Emery, 1906
12) P. nitidula Emery, 1888
13) P. indica Mayr, 1879

14) P. obscurior Forel, 1886

A remark about synonymy

Anergates friedlandi Creighton, 1934 is now a synonym from Anergates atratulus (Schenck, 1852), more precisely an introduced form in North Americe. So, the name is Anergates atratulus (Schenck, 1852) or, if you follow Ward, Brady, Fisher, Schultz, 2015 ("2014"), Tetramorium atratulum (Schenck, 1852)...

Synonyms of host species

- T. caespitum (Linnaeus, 1758) (= Formica caespitum Linnaeus, 1758)
- T. impurum (Förster, 1850) (= Myrmica impura Förster, 1850)
- P. megacephala (Fabricius, 1793) (= Formica megecephala Fabricius, 1793)
- P. megacephala (Fabricius, 1793) subsp. melancholica Santschi, 1912 was originally described as P. punctulata Mayr, 1866 st. melancholica Santschi, 1912
- The host species P. strobeli Emery, 1906 was known, during much of its history, as a subspecies of an other species, nl. P. nitidula Emery, 1888 subsp. richteri Forel, 1909 (a much used junior synonym of the older P. perversa Forel, 1908 subsp. richteri Forel, 1909) and known, in 1922, for a while as P. strobeli Emery, 1906 subsp. richteri Forel, 1909, this since Santschi, 1916. The species P. strobeli Emery, 1906 was revived from synonymy (with P. nitidula Emery, 1888 since Wilson, 2003) and the subspecies was synonymized with it in Casadei-Ferreira, Economo, Feitosa, 2020.
- P. obscurior Forel, 1886 was revived from synonymy with the 1 page older P. susannae Forel, 1886 and this in Casadei-Ferreira, Economo, Feitosa, 2020.
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Sonntag 28. November 2021, 20:34

For the rest a few changes in literature lists.

I removed Passera from the list "not in the lists" to the "Teleutomyrmex list", so it includes now 103 references.

I changed the year in one of the "Teleutomyrmex list" references to the correct year, nl. 2006 i.s.o. 2005.

I included four references in the "not in the lists", nl. two Dutch and two French, all from Belgium.

I did put in the page numbers in the Huber reference.
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Sonntag 28. November 2021, 20:35

Official literature lists 54.pdf
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Sonntag 28. November 2021, 20:53

An official literature list of Teleutomyrmex Kutter, 1950.

Baroni Urbani, C., 1967, “Le distribuzioni geografiche discontinue dei Formicidi mirmecobiotici.” Archivio Botanico e Biogeografico Italiano, vol. 43, [=(4)12], p. 355-365.

Baur, A., Buschinger, A., Zimmermann, F. K., 1993, “Molecular cloning and sequencing of 18S rDNA gene fragments from six different ant species.” Insectes Sociaux, vol. 40, p. 325-335.

Baur, A., Chalwatzis, N., Buschinger, A., Zimmermann, F. K., 1995, “Mitochondrial DNA sequences reveal close relationships between social parasitic ants and their host species.” Current Genetics, vol. 28, p. 242-247.

Baur, A., Sanetra, M., Chalwatzis, N., Buschinger, A., Zimmermann, F. K., 1996, “Sequence comparisons of the internal transcribed spacer region of ribosomal genes support close relationships between parasitic ants and their respective host species (Hymenoptera: Formicidae).” Insectes Sociaux, vol. 43, p. 53-67.

Bernard, F., 1967 ["1968"], “Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale.” Paris, Masson, 411 pp.

Blaimer, B. B., Ward, P. S., Schultz, T. R., Fisher, B. L., Brady, S. G., 2018, “Paleotropical Diversification Dominates the Evolution of the Hyperdiverse Ant Tribe Crematogastrini (Hymenoptera: Formicidae).” Insect Systematics and Diversity, vol. 2, no. 5, article no. 3, p. 1-14 (+ 2 supporting files.).

Bolton, B., 1976, “The ant tribe Tetramoriini (Hymenoptera: Formicidae). Constituent genera, review of smaller genera and revision of Triglyphothrix Forel.” Bulletin of the British Museum (Natural History) (Entomology), vol. 34, p. 281-379.

Bolton, B., 1994, “Identification guide to the ant genera of the world.” Cambridge, Mass., Harvard University Press, 222 pp.

Bolton, B., 1995, “A taxonomic and zoogeographical census of the extant ant taxa (Hymenoptera: Formicidae).” Journal of Natural History, vol. 29, p. 1037-1056.

Bolton, B., 1995, “A new general catalogue of the ants of the world.” Cambridge, Mass., Harvard University Press, 504 pp.

Bolton, B., 2003, “Synopsis and Classification of Formicidae.” Memoirs of the American Entomological Institute, vol. 71, 370 pp.

Borowiec, L., 2014, “Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae).” Genus (Wroclaw), vol. 25, no. 1-2, p. 1-340.

Bourke, A. F. G., Franks, N. R., 1991, “Alternative adaptations, sympatric speciation and the evolution of parasitic, inquiline ants.” Biological Journal of the Linnean Society, vol. 43, no. 3, p. 157-178.

Brown, W. L., Jr., 1973, “A comparison of the Hylean and Congo-West African rain forest ant faunas.” P. 161-185 in: Meggers, B. J., Ayensu, E. S., Duckworth, W. D. (editors.), 1973, “Tropical forest ecosystems in Africa and South America: a comparative review.” Smithsonian Institution Press, Washington, DC., viii + 350 pp.

Brun, R., 1952, “Das zentralnervensystem von Teleutomyrmex schneideri Kutt. (Hym. Formicid.). III. Mitteilung.” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 25, no. 2, p. 73-86.

Brun, R., 1959, “Le cerveau des fourmis et des insectes en général comme instrument de formation des réflexes conditionnés.” P. 11-25 in: International Union of Biological Sciences, Union Internationale des Sciences Biologiques, Experimental Psychology and Animal Behaviour Section, 1959, “Animal psychology seminars: Strasbourg University, October 1956, and Brussels University, August 1957.” London, Pergamon Press, 148 pp.

Buschinger, A., 1970, “Neue Vorstellungen zur Evolution des Sozialparasitismus und der Dulosis bei Ameisen (Hym., Formicidae).” Biologisches Zentralblatt, vol. 89, no. 3, p. 273-299.

Buschinger, A., 1971, “Zur Verbreitung und Lebensweise sozialparasitischer Ameisen des Schweizer Wallis (Hym., Formicidae).” Zoologischer Anzeiger, vol. 186, no. 1/2, p. 47-59.

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-> Dobrzanska, J., Dobrzanski, J., 1988, “1. Inkwilinizm.”, or “[Inquilinism.]” Kosmos, vol. 37, no. 4, p. 617-629.
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- His name is spelled “Göszwald” in this publication!

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An original “Book Review”…

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Schenck, C. F., 1853b, “Die nassauischen Ameisen-Species. (Fortsetzung.).” Stettiner Entomologische Zeitung, vol. 14, p. 185-198.

van Boven, J. K. A., 1977, “De mierenfauna van België (Hymenoptera: Formicidae).” Acta Zoologica et Pathologica Antverpiensia, vol. 67, p. 1-191.

Wagner, H. C., Arthofer, W., Seifert, B., Muster, C., Steiner, F. M., Schlick-Steiner, B. C., 2017, “Light at the end of the tunnel: Integrative taxonomy delimits cryptic species in the Tetramorium caespitum complex (Hymenoptera: Formicidae).” Myrmecological News, vol. 25, p. 95–129 (+ 6 supporting files.).

Ward, P. S., Brady, S. G., Fisher, B. L., Schultz, T. R., 2015 (“2014”), “The evolution of myrmicine ants: Phylogeny and biogeography of a hyperdiverse ant clade (Hymenoptera: Formicidae).” Systematic Entomology, vol. 40, no. 1, p. 61-81 (+ 9 supporting files.). (Article first published online: 23 July 2014).

Wheeler, G. C., Wheeler, J., 1955c, “The ant larvae of the myrmicine tribe Solenopsidini.” American Midland Naturalist, vol. 54, p. 119-141.

Wheeler, W. M., 1909g, “Observations on some European ants.” Journal of the New York Entomological Society, vol. 17, p. 172-187.

Schlick-Steiner, B. C., Steiner, F. M., Sanetra, M., Heller, G., Stauffer, C., Christian, E., Seifert, B., 2005, “Queen size dimorphism in the ant Tetramorium moravicum (Hymenoptera, Formicidae): morphometric, molecular genetic and experimental evidence.” Insectes Sociaux, vol. 52, p. 186-193.

Schulz, A., 1996, “Tetramorium rhenanum nov. spec. vom "Mittleren Rheintal" in Deutschland (Hymenoptera: Formicidae).” Linzer Biologische Beiträge, vol. 28, p. 391-412.

21 + 2 references.

And a few references missed in the literature lists of Teleutomyrmex Kutter, 1950 and Anergates Forel, 1874.

Agosti, D., Majer, J. D., Alonso, L. E., Schultz, T. R. (editors.), 2000, “Ants: Standard Methods for Measuring and Monitoring Biodiversity.” Washington, Smithsonian Institution Press, xix + 280 pp.

Agosti, D., Majer, J. D., Alonso, L. E., Schultz, T. R. (editors.), 2000, “Sampling ground-dwelling ants: case studies from the world's rain forests.” Perth, Australia, Curtin University School of Environmental Biology (Bulletin No. 18), xii + 75 pp.

Boer, P., 2010, “Mieren van de Benelux.” ′s-Graveland, Netherlands, Stichting Jeugdbondsuitgeverij, 184 pp.

Boer, P., 2015, “Mieren van de Benelux. Tweede - herziene druk.” ′s-Graveland, Netherlands, Stichting Jeugdbondsuitgeverij, 184 pp.

Bolton, B., Collingwood, C. A., 1975, “Handbooks for the identification of British Insects. Hymenoptera, Formicidae.” Handbooks for the Identification of British Insects, vol. 6, no. 3(c), p. 1-34.

Bourke, A. F. G., Franks, N. R., 1995, “Social Evolution in Ants.” Princeton, Princeton University Press, xiii + 529 pp.

Buschinger, A., 1985, “Staatenbildung der Insekten.” Erträge der Forschung, Band 223. Darmstadt, Wissenschaftliche Buchgesellschaft, x + 211 pp.

Donisthorpe, H., 1927, “British ants, their life history and classification (2nd edn.).” London, G. Routledge and Sons, xvi + 436 pp.

Donisthorpe, H., 1927, “The guests of British ants, their habits and life-histories.” London, G. Routledge and Sons, xxii + 244 pp.

Emery, C., 1910, “Hymenoptera. Fam. Formicidae. Subfam. Dorylinae.” Genera Insectorum, vol. 102, p. 1-34.

Emery, C., 1911, “Hymenoptera. Fam. Formicidae. Subfam. Ponerinae.” Genera Insectorum, vol. 118, p. 1-125.

Emery, C., 1913 ("1912"), “Hymenoptera. Fam. Formicidae. Subfam. Dolichoderinae.” Genera Insectorum, vol. 137, p. 1-50.

Emery, C., 1921, “Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. [part].” Genera Insectorum, vol. 174A, p. 1-94 (+ 7 plates.).

Emery, C., 1922, “Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. [part].” Genera Insectorum, vol. 174B, p. 95-206.

Emery, C., 1924 ("1922"), “Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. [concl.].” Genera Insectorum, vol. 174C, p. 207-397.

Emery, C., 1925, “Hymenoptera. Fam. Formicidae. Subfam. Formicinae.” Genera Insectorum, vol. 183, p. 1-302.

Forel, A., 1915, “Fauna insectorum helvetiae. Hymenoptera. Formicidae. Die Ameisen der Schweiz.” Mitteilungen der Schweizerischen Entomologischen Gesellschaft, vol. 12 (Beilage), p. 1-77.

Forel, A,. 1920, “Les fourmis de la Suisse. Notices anatomiques et physiologiques, architecture, distribution géographique, nouvelles expériences et observations de moeurs. Seconde édition revue et corrigée.” La Chaux-de-Fonds, Imprimerie Coopérative, xvi + 333 pp.
- Without the section “Systématique”, so without descriptions et al.

Forel, A., 1921, “Le monde social des fourmis comparé à celui de l'homme. Tome 1. Genèse, formes, anatomie, classification. géographie, fossiles.” Genève, Librairie Kundig, xiv + 192 pp.

Forel, A., 1921, “Le monde social des fourmis comparé à celui de l'homme. Tome 2. Sensations, physiologie, fourmis et plantes, hôtes, parasites, nids.” Genève, Librairie Kundig, iii + 184 pp.

Forel, A., 1922, “Le monde social des fourmis comparé à celui de l'homme. Tome 3. Appareils d'observation. Fondation des foumilières. Moeurs à l'intérieur des nids. Bétail, jardins, fourmis parasites. (Avec appendice du Dr. E. Bugnion.).” Genève, Librairie Kundig, vii + 227 pp.

Forel, A., 1923, “Le monde social des fourmis comparé à celui de l'homme. Tome 4. Alliances et guerres, parabiose, lestobiose, esclavagisme.” Genève, Librairie Kundig, vii + 172 pp.

Forel, A., 1923, “Le monde social des fourmis comparé à celui de l'homme. Tome 5. Moeurs specialisées. Epilogue, les fourmis. Les termites et l'homme.” Genève, Librairie Kundig, vi + 174 pp.

Forel, A., 1948, “Die Welt der Ameisen. (Ausgewählt und übersetzt von Heinrich Kutter.).” Zürich, Rotapfel, ?? + 275 pp.

Gösswald, K., 1989, “Die Waldameise, Band 1: Biologische Grundlagen, Ökologie und Verhalten.” Wiesbaden, AULA-Verlag, xi + 660 pp.

Gösswald, K., 1990, “Die Waldameise, Band 2: Die Waldameise im Ökosystem Wald, ihr Nutzen und ihre Hege.” Wiesbaden, AULA-Verlag, x + 510 pp.

Gotwald, W. H., Jr., 1995, “Army ants: the biology of social predation.” Ithaca, New York, Cornell University Press, xviii + 302 pp.

Gould, W., 1747, “An Account Of English Ants.” London, A. Millar, xvi + 112 pp.

Hermann, H. R. (editor.), 1979, “Social Insects, Vol. 1.” New York, Academic Press, xv + 437 pp.

Hermann, H. R. (editor.), 1981, “Social Insects, Vol. 2.” New York, Academic Press, xiii + 491 pp.

Hermann, H. R. (editor.), 1982, “Social Insects, Vol. 3.” New York, Academic Press, xiii + 459 pp.

Hermann, H. R. (editor.), 1982, “Social Insects, Vol. 4.” New York, Academic Press, ?? + 385 pp.

Hermann, H. R. (editor.), 1984, “Defensive Mechanisms in Social Insects.” New York, Praeger Publishers, xii + 259 pp.

Hölldobler, B. K., Wilson, E. O., 2008, “The Superorganism. The Beauty, Elegance, and Strangeness of Insect Societies.” New York, W. W. Norton & Company, xxi + 522 pp.

Hölldobler, B. K., Wilson, E. O., 2011, “The Leafcutter Ants. Civilisation by Instinct.” New York, W. W. Norton & Company, 160 pp.

Kronauer, D. J. C., 2020, “Army Ants: Nature’s Ultimate Social Hunters.” Cambridge, Mass., Harvard University Press, ?? + 384 pp.

Künckel d’Herculais, J., 1882, “Les Merveilles de la Nature. L’Homme et les Animaux. A. E. Brehm. Merveilles de la Nature. Les Insectes. Les Myriopodes, les Arachnides et les Crustacés. Édition Française par J. Künckel d’Herculais, aide-naturaliste au Muséum d’Histoire Naturelle. Tome 1.” Paris, Librairie J.-B. Baillière et Fils, viii + 720 pp.

Künckel d’Herculais, J., 1882, “Les Merveilles de la Nature. Les Insectes. A. E. Brehm. Merveilles de la Nature. Les Insectes. Les Myriopodes, les Arachnides. Édition Française par J. Künckel d’Herculais, aide-naturaliste au Muséum d’Histoire Naturelle. Tome 2.” Paris, Librairie J.-B. Baillière et Fils, iv + 802 pp.
- Tome 2 starts with “Les Formicides – Formicidae
Die Ameisen. – The Ants.” on p. 1-150.

Kutter, H., 1920, “”Gehe hin zur Ameise!” Anleitung zur selbständigen Ameisenforschung.” Naturwissenschaftliche Beobachtungsbücher, Band 1-2. Bern und Leipzig, Ernst Bircher Verlag, viii + 165 pp.

Lach, L., Parr, C. L., Abbott, K. (editors.), 2009, “Ant ecology.” Oxford, Oxford University Press, xvii + 410 pp.

Maeterlinck, M., 1930, “La Vie des Fourmis.” Bibliothèque Charpentier. Paris, Eugène Fasquelle, éditeur, 254 pp.

Schmitz, H., 1906, “Das Leben der Ameisen und ihrer Gäste.” Regensburg, G. J. Manz, 190 pp.

Schmitz, H., 1915, “De Nederlandsche mieren en haar gasten.” Maastricht, C. Goffin, iv + 146 pp.

Schneirla, T. C., 1971, “Army ants. A study in social organization. (Edited by H. R. Topoff.).” San Francisco, W. H. Freeman & Co., xx + 349 pp.

Senden, L., 1929, “Uit het huishouden der mieren. Deel 1.” Davidsfonds. Nr. 224. Brugge, Drukk. Excelsior, 182 pp.

Senden, L., 1929, “Uit het huishouden der mieren. Deel 2.” Davirsfonds. Nr. 225. Brugge, Drukk. Excelsior, 136 pp.

Skinner, G. J., 1987, “Ants of the British Isles.” Shire Natural History Series, vol. 21, p. 1-24.

Skinner, G. J., Allen, G. W., 1996, “Ants.” Naturalists' Handbook, vol. 24, Slough, The Richmond Publishing Co. Ltd., ii + 83 pp.
- This volume, in its introduction, speaks, as one of only a few, about Solenopsis wagneri Santschi, 1916e.

Stitz, H., 1939, “Die Tierwelt Deutschlands und der angrenzenden Meersteile nach ihren Merkmalen und nach ihrer Lebensweise. 37. Theil. Hautflüger oder Hymenoptera. I: Ameisen oder Formicidae.” Jena, G. Fischer, 428 pp.

Taber, S. W., 2000, “Fire Ants.” Texas A&M University Press, 328 pp.

Vander Meer, R. K., Jaffe, K., Cedeno, A. (editors.), 1990, “Applied myrmecology: a world perspective.” Boulder, Westview Press, xv + 741 pp.

Weber, N. A., 1972, “Gardening ants, the attines.” Philadelphia, American Philosophical Society, xvii + 146 pp.

Verhaeghe, J. C., Deligne, J., De Vos, L., Quinet, W., 1984, “Les Cahiers du Viroin, 2. Les fourmis de nos régions. Introduction à la biologie sociale des fourmis.” Triegnes, Editions D.I.R.E., Centre Paul Brien, 152 pp.

Wheeler, W. M., 1910, “Ants: Their Structure, Development and Behavior.” New York, Columbia University Press, xxv + 663 pp.

Wheeler, W. M., 1923, “Social life among the insects.” New York, Harcourt, Brace and Co., vii + 375 pp.

Wheeler, W. M., 1926, “The natural history of ants. From an unpublished manuscript in the archives of the Academy of Sciences of Paris by René Antoine Ferchault de Réaumur. Translated and annotated by William Morton Wheeler.” New York, A. Knopf, 280 pp.

Wheeler, W. M., 1928, “The social insects: their origin and evolution.” New York, Harcourt, Brace and Co., xviii + 378 pp.

Williams, D. F. (editor.), 1994, “Exotic ants. Biology, impact, and control of introduced species.” Boulder, Westview Press, xvii + 332 pp.

Wilson, E. O., 1990, “Excellence in Ecology, Vol. 2: Success and Dominance in Ecosystems: The Case of the Social Insects.” Oldendorf/Luhe, Ecology Institute, xxi + 104 pp.

59 references.

The beginning of modern myrmecology.

Huber, P., 1810, “Recherches sur les mœurs des fourmis indigènes.” Paris, Chez J. J. Paschoud, xvi + 328 pp. (+ 2 plates!)

Latreille, P. A., 1798, “Essai sur l'histoire des fourmis de la France.” Brive, F. Bourdeaux, 50 pp.

A few marvellous biographies.

Carpenter, H., 1977, “J. R. R. Tolkien: A Biography.” London, George Allen & Unwin, ?? pp.

Evans, M. A., Evans, H. E., 1970, “William Morton Wheeler, biologist.” Cambridge, Mass., Harvard University Press, 363 pp.

Pais, A., 1982, “'Subtle is the Lord ...': The Science and the Life of Albert Einstein.” Oxford, Oxford University Press, ?? pp.

Pais, A., 1994, “Einstein Lived Here.” Oxford, Clarendon Press/Oxford University Press, ?? pp.

And then, of course, there is the biography of Cristoforo Colombo/Christopher Columbus, somewhere in the late eighties or in the beginning of the nineties, but for the moment I can’t remember more about it…

Two books NOT to have…

Gösswald, K., 2012, “Die Waldameise. Biologie, Ökologie und forstliche Nutzung.” 1. Auflage, gekürzte Sonderausgabe in einem Band, Aula Verlag, Wiebelsheim, xii + 630 pp. (Editors unknown!).

Tschinkel, W. R., 2006, “The Fire Ants.” Belknap Press of Harvard University Press, 744 pp.

2 + 5 + 2 references.

Literature list of important Solenopsis-references.

Santschi, F., 1916, “Formicides sudaméricains nouveaux ou peu connus.” Physis, vol. 2, p. 365-399.

Santschi, F., 1923, “Solenopsis et autres fourmis néotropicales.” Revue Suisse de Zoologie, vol. 30, no. 8, p. 245-273.

Creighton, W. S., 1930, “The New World species of the genus Solenopsis.” Proceedings of the American Academy of Arts and Sciences, vol. 66, no. 2, p. 39-151 (+ 8 plates!).

Creighton, W. S., 1950, “The ants of North America.” Bulletin of the Museum of Comparative Zoology [at Harvard College], vol. 104, p. 1-585 (+ 57 plates!).

Wilson, E. O., 1951, “Variation and adaptation in the imported fire ant.” Evolution, vol. 5, no. 1, p. 68-79.

Wilson, E. O., 1952, “O complexo Solenopsis saevissima na América do Sul (Hymenoptera: Formicidae).” Memórias do Instituto Oswaldo Cruz. Rio de Janeiro, vol. 50, p. 49-59.

Wilson, E. O., 1952, “The Solenopsis saevissima complex in South America (Hymenoptera: Formicidae).” Memórias do Instituto Oswaldo Cruz, Rio de Janeiro, vol. 50, p. 60-68.
- Translation of preceding.

Wilson, E. O., 1953, “Origin of the variation in the imported fire ant.” Evolution, vol. 7, no. 3, p. 262-263.

Wheeler, G. C., Wheeler, J., 1955, “The ant larvae of the myrmicine tribe Solenopsidini.” American Midland Naturalist, vol. 54, no.1, p. 119-141.

Wilson, E. O., Brown, W. L., Jr., 1958, “Recent changes in the introduced population of the fire ant Solenopsis saevissima (Fr. Smith).” Evolution, vol. 12, no. 2, p. 211-218.

Snelling, R. R., 1963, “The United States species of fire ants of the genus Solenopsis, subgenus Solenopsis Westwood, with synonymy of Solenopsis aurea Wheeler (Hymenoptera: Formicidae).” California Department of Agriculture, Bureau of Entomology, Occasional Papers, no. 3, p. 1-11 (+ 2 plates!).

Ettershank, G., 1966, “A generic revision of the world Myrmicinae related to Solenopsis and Pheidologeton (Hymenoptera: Formicidae).” Australian Journal of Zoology, vol. 14, p. 73-171.

Buren, W. F., 1972, “Revisionary studies of the taxonomy of the imported fire ants.” Journal of the Georgia Entomological Society, vol. 7, p. 1-26.

O’Neal, J., Markin, G. P., 1975, “The larval instars of the imported fire ant, Solenopsis invicta Buren.” Journal of the Kansas Entomological Society, vol. 48, p. 141-151.

Wheeler, G. C., Wheeler, J., 1977, “Supplementary studies on ant larvae: Myrmicinae.” Transactions of the American Entomological Society, vol. 103, no. 1450, p. 581-602.

Petralia, R. S., Vinson, S. B., 1979, “Developmental morphology of larvae and eggs of the imported fire ant, Solenopsis invicta.” Annals of the Entomological Society of America, vol. 72, p. 472–484.

Bolton, B., 1987, “A review of the Solenopsis genus-group and revision of Afrotropical Monomorium Mayr (Hymenoptera: Formicidae).” Bulletin of the British Museum (Natural History), Entomology, vol. 54, no. 3, p. 263-452.

Carlton, C. E., 1987, “Identification of Arkansas fire ant (Hymenoptera: Formicidae: Solenopsis spp.) workers.” Report Series, University of Arkansas Agricultural Experiment Station, no. 301, p. 1-9 (= 14pp.).

Trager, J. C., 1991, “A revision of the fire ants, Solenopsis geminate group (Hymenoptera: Formicidae: Myrmicinae).” Journal of the New York Entomological Society, vol. 99, no.2, p. 141-198.

Taber, S. W., 2000, “Fire Ants.” Texas A&M University Press, 328 pp.

Pitts, J. P., 2002, “A cladistic analysis of the Solenopsis saevissima species-group (Hymenoptera: Formicidae).” Dissertation, University of Georgia, vi + 266 pp.

Tschinkel, W. R., Mikheyev, A. S., Storz, S. R., 2003, “Allometry of workers of the fire ant, Solenopsis invicta.” Journal of Insect Science, vol. 3, no.2, p. 1-11.

Pitts, J. P., McHugh, J. V., Ross, K. G., 2005, “Cladistic analysis of the fire ants of the Solenopsis saevissima species-group (Hymenoptera: Formicidae).” Zoologica Scripta, vol. 34, no. 5, p. 493-505.

Tschinkel, W. R., 2006, “The Fire Ants.” Belknap Press of Harvard University Press, 744 pp.

Araujo, M., Tschinkel, W. R., 2010, “Worker allometry in relation to colony size and social form in the fire ant, Solenopsis Invicta.” Journal of Insect Science, vol. 10, no. 94, p. 1-10.

Tschinkel, W.R., 2013, “The morphometry of Solenopsis fire ants.” PlosOne, vol. 8, no. 11, p. 1-22 (+ 1 supporting file.).

Chialvo, P., 2017, “Species delimitation of the native North American fire ants (Formicidae: Solenopsis): results and applications to life sciences education.” Dissertation, University of Georgia, xi + 124 pp.

Chialvo, P., Gotzek, D. A., Shoemaker, D., Ross, K. G., 2018 (“2017”), “Genetic analyses reveal cryptic diversity in the native North American fire ants (Hymenoptera: Formicidae: Solenopsis).” Systematic Entomology, vol. 43, no. 1, p. 109-122 (+ 2 supporting files.).

Pitts, J. P., Camacho, G. P., Gotzek, D., McHugh, J. V., Ross, K. G., 2018, “Revision of the fire ants of the Solenopsis saevissima species-group (Hymenoptera: Formicidae).” Proceedings of the Entomological Society of Washington, vol. 120, no.2, p. 308–411 (+ 4 supporting files.).

29 references.

My personal introduction books to ants...

The most important books that deal only with ants (in general!):

01) Hölldobler, B. K., Wilson, E. O., 1994, “Journey to the Ants: A Story of Scientific Exploration.” Cambridge, Mass., Belknap Press of Harvard University Press, 304 pp.

02) Hölldobler, B. K., Wilson, E. O., 1990, “The Ants.” Cambridge, Mass., Belknap Press of Harvard University Press, xiv + 732 pp.

03) Hölldobler, B. K., Wilson, E. O., 2011, “The Leafcutter Ants. Civilisation by Instinct.” New York, W. W. Norton & Company, 160 pp.

04) Bolton, B., 1994, “Identification guide to the ant genera of the world.” Cambridge, Mass., Harvard University Press, 222 pp.

05) Bolton, B., 2003, “Synopsis and Classification of Formicidae.” Memoirs of the American Entomological Institute, vol. 71, 370 pp.

06) Wheeler, W. M., 1910, “Ants: Their Structure, Development and Behavior.” New York, Columbia University Press, xxv + 663 pp.

07) Agosti, D., Majer, J. D., Alonso, L. E., Schultz, T. R. (editors.), 2000, “Ants: Standard Methods for Measuring and Monitoring Biodiversity.” Washington, Smithsonian Institution Press, xix + 280 pp.

08) Bourke, A. F. G., Franks, N. R., 1995, “Social Evolution in Ants.” Princeton, Princeton University Press, xiii + 529 pp.

09) Dumpert, K., 1978, “Das Sozialleben der Ameisen.” (First Edition.). Pareys Studientexte, nr. 18. Mit 95 Abbildungen. Verlag Paul Parey, Berlin und Hamburg, 1977, 253 pp.

10) Dumpert, K., 1994, “Das Sozialleben der Ameisen. 2., neubearbeitete Auflage.” (Second Edition.). Pareys Studientexte, nr. 18. Mit 94 Abbildungen. Verlag Paul Parey, Berlin und Hamburg, 1994, 257 pp.

A small selection of books on social insects that are also very good on ants:

01) Wilson, E. O., 1971, “The insect societies.” Cambridge, Mass., Harvard University Press, x + 548 pp.

02) Hermann, H. R. (editor.), 1979, “Social Insects, Vol. 1.” New York, Academic Press, xv + 437 pp.

03) Hermann, H. R. (editor.), 1981, “Social Insects, Vol. 2.” New York, Academic Press, xiii + 491 pp.

04) Hermann, H. R. (editor.), 1982, “Social Insects, Vol. 3.” New York, Academic Press, xiii + 459 pp.

05) Hermann, H. R. (editor.), 1982, “Social Insects, Vol. 4.” New York, Academic Press, ?? + 385 pp.

06) Hermann, H. R. (editor.), 1984, “Defensive Mechanisms in Social Insects.” New York, Praeger Publishers, xii + 259 pp.

07) Wilson, E. O., 1990, “Excellence in Ecology, Vol. 2: Success and Dominance in Ecosystems: The Case of the Social Insects.” Oldendorf/Luhe, Ecology Institute, xxi + 104 pp.

08) Hölldobler, B. K., Wilson, E. O., 2008, “The Superorganism. The Beauty, Elegance, and Strangeness of Insect Societies.” New York, W. W. Norton & Company, xxi + 522 pp.

There are still a lot more important books like e.g. for ants:

01) Passera, L., 1984, “L'Organisation Sociale des Fourmis.” Toulouse, Editions Privat, 360 pp.

02) Gösswald, K., 1985, “Organisation und Leben der Ameisen.” Stuttgart, Wissenschaftliche Verlagsgesellschaft mbH, 355 pp.

03) Gösswald, K., 1989, “Die Waldameise, Band 1: Biologische Grundlagen, Ökologie und Verhalten.” Wiesbaden, AULA-Verlag, xi + 660 pp.

04) Gösswald, K., 1990, “Die Waldameise, Band 2: Die Waldameise im Ökosystem Wald, ihr Nutzen und ihre Hege.” Wiesbaden, AULA-Verlag, x + 510 pp.

And many more...

A few determination books from one author…

01) Seifert, B., 1996, “Ameisen: beobachten, bestimmen.” Naturbuch Verlag, Augsburg, 351 pp.

02) Seifert, B., 2007, “Die Ameisen Mittel- und Nordeuropas.” Lutra Verlags- und Vertriebsgesellschaft, Tauer/Klitten, 368 pp.

03) Seifert, B., 2018, “The ants of Central and North Europe.” Lutra Verlags- und Vertriebsgesellschaft, Tauer/Boxberg, 407 pp.

And one of the many important articles:

Joshua Blu Buhs, 2000, "Building on Bedrock: William Steel Creighton and the Reformation of Ant Systematics, 1925-1970." Journal of the History of Biology, vol. 33, no. 1, p. 27-70.

25 books and 1 article.
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Samstag 4. Dezember 2021, 18:51

“Emery’s Rule” in its strict and its loose version.

In 1909, the entomologist/taxonomist Carlo Emery made an important generalization and noted that social parasites among insects and their hosts share common ancestry and hence tend to be parasites of species or genera to which they are closely related. Over time, this pattern has been recognized in many additional cases, and generalized to what is now known as “Emery's rule”. The pattern is best known for various taxa of Hymenoptera. The significance and general relevance of this pattern are still a matter of some debate, as a great many exceptions exist, though a common explanation for the phenomenon when it occurs is that the parasites may have started as facultative parasites within the host species itself (such forms of intraspecific parasitism are well-known.), but later became reproductively isolated and split off from the ancestral species, a form of sympatric speciation.

In the “strict version” of Emery’s rule, social parasitic species are their host’s closest relatives (a sister taxon to its host in a phylogenetic sense.), and likely evolved from their host’s lineage by sympatric speciation, or through a combination of allopatric and subsequent sympatric speciation. In the “loose version” of Emery’s rule, social parasites are close relatives of their host, but not sister species. Although few host-parasite pairs have been subjected to molecular phylogenetic analysis, studies to date support at least the loose version of Emery’s rule.

The “Kutter-Wilson Paradox”.

There are only about 230 known parasitic species of ants among the 12,500 or so described ant species. Despite their rarity, they are common in a few subfamilies like the Myrmicinae and the Formicinae, and common in temperate ants but rare in tropical ants. Why is there such a strong bias in both the taxonomic and ecological distributions of social parasites and their hosts, the “Kutter-Wilson Paradox”? The selective forces and ecological conditions that favour social parasitism continue to be researched and discussed. Factors like cooler temperatures and polygyny (i.e., colonies with multiple queens.) are important considerations, but cannot explain the evolution of social parasitism in all cases.

The “Inquiline Syndrome”.

Wilson, E. O. [“The insect societies”, 1971, p. 374] and Hölldobler, B. K. and Wilson, E. O. [“The ants”, 1990, p. 467] defined the “Inquiline Syndrome” and listed 41 characters of morphological, behavioural and life history traits that evolved convergently in inquiline social parasites (a 19 points of special interesses-list.). Not all those inquilines have all the characters but they have most of them. The ants of the genus Teleutomyrmex display 36 characters of the list.

Reference:

Emery, C., 1909e, “Über den Ursprung der dulotischen, parasitischen und myrmekophilen Ameisen.” Biologisches Centralblatt, vol. 29, p. 352-362.
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Sonntag 5. Dezember 2021, 18:09

A new host and observations about copulation and adoption in Tetramorium-Strongylognathus nests.

Purkart, A., Wagner, H.CPurkart, A., Wagner, H.C., Goffová, K., Selnekovič, D., Holecová, M. 2021. Laboratory observations on Anergates atratulus (Schenck, 1852): mating behaviour, incorporation into host colonies, and competition with Strongylognathus testaceus (Schenck, 1852). Biologia.,

https://www.antwiki.org/wiki/images/e/e ... 1-y%29.pdf
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Sonntag 5. Dezember 2021, 18:12

The new host of Anergates is:

T. staerckei Kratochvíl, in Kratochvíl, Novák, Šnoflák, 1944 (= Tetramorium caespitum (Linnaeus, 1758) subsp. hungarica Röszler, 1935 ("1933-34") var. staerckei Röszler, 1936)
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Sonntag 5. Dezember 2021, 18:13

With the new host...

The extreme, workerless inquilines 56.pdf
(210.47 KiB) 1086-mal heruntergeladen
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Dienstag 14. Dezember 2021, 10:04

A review of the last article:

viewtopic.php?f=23&t=367&start=400#p22654
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Freitag 17. Dezember 2021, 21:50

The extreme, workerless inquilines.

Teleutomyrmex Kutter, 1950

01) Teleutomyrmex schneideri Kutter, 1950
(= Tetramorium schneideri (Kutter, 1950), by Ward, Brady, Fisher, Schultz, 2015 ("2014"))
(not Tetramorium schneideri Emery, 1898)
(= Tetramorium inquilinum Ward, Brady, Fisher, Schultz, 2015 ("2014"))
02) Teleutomyrmex kutteri Tinaut, 1990
(= Tetramorium kutteri (Tinaut, 1990), by Ward, Brady, Fisher, Schultz, 2015 ("2014"))
(not Tetramorium semilaeve André, 1883 var./subsp. kutteri Santschi, 1927)
03) Teleutomyrmex seiferti Kiran & Karaman, in Kiran, Karaman, Lapeva-Gjonova, Aksoy, 2017
(= Tetramorium seiferti (Kiran & Karaman, in Kiran, Karaman, Lapeva-Gjonova, Aksoy, 2017), by analogy)
04) Teleutomyrmex buschingeri Lapeva-Gjonova, in Kiran, Karaman, Lapeva-Gjonova, Aksoy, 2017
(= Tetramorium buschingeri (Lapeva-Gjonova, in Kiran, Karaman, Lapeva-Gjonova, Aksoy, 2017), by analogy)

Not yet described species of extreme, workerless inquiline, from the genus Teleutomyrmex Kutter, 1950

05) The new, undescribed species from Teleutomyrmex Kutter, 1950 from Farab, Turkmenistan… See Dlussky, Soyunov, Zabelin, 1990 [“1989”].

Anergates Forel, 1874

06) Anergates atratulus (Schenck, 1852), by Forel, 1874
(= Myrmica atratula Schenck, 1852)
[Also described as new by Schenck, 1853b]
(= Tetramorium atratulum (Schenck, 1852), by Mayr, 1855)
[= Tomognathus atratulus (Schenck, 1852), by Mayr, 1863 following Mayr, 1861, obsolete combination.]
(= Tetramorium atratulum (Schenck, 1852), by Ward, Brady, Fisher, Schultz, 2015 ("2014"))
07) Anergates friedlandi Creighton, 1934
[= Tetramorium friedlandi (Creighton, 1934), by analogy]

Tetramorium Mayr, 1855
(Only a few species in a big genus.)

08) Tetramorium microgyna Santschi, 1918
09) Tetramorium parasiticum Bolton, 1980

Pheidole Westwood, 1839
(Only a few species in a big genus.)

10) Pheidole neokohli Wilson, 1984
(= Pheidole kohli (Wasmann, 1915), by Wilson, 1984)
(= Anergatides kohli Wasmann, 1915)
(not Pheidole kohli Mayr, 1901)
11) Pheidole acutidens (Santschi, 1922), by Wilson, 1984
(= Bruchomyrma acutidens Santschi, 1922)
12) Pheidole argentina (Bruch, 1932), by Wilson, 1984
(= Gallardomyrma argentina Bruch, 1932)
13) Pheidole parasitica Wilson, 1984

Excluded from the extreme, workerless inquilines
Once this species was included in the extreme, workerless inquilines but now it is considered to be a workerless inquiline without extreme reductions, e.g. no pupoid males but normal ones. The decision to exclude it was made by Edward Osborne Wilson in 1984 in a study of the inquilines in the genus Pheidole Westwood, 1839.

Pheidole Westwood, 1839
(Only one species in a big genus.)

14) Pheidole kusnezovi Wilson, 2003
(= Pheidole symbiotica (Kusnezov, 1952), by Wilson, 1984)
(= Eriopheidole symbiotica Kusnezov, 1952)
(not Pheidole symbiotica Wasmann, 1909)

Distribution

01) Europe (Alps, Pyrenees and Northern Spain)
02) Europe (Southern Iberia)
03) Turkey (Anatolia)
04) Europe (Southern Balkans)

05) Turkmenistan

06) Europe
07) North America

08) Southern Africa
09) Southern Africa

10) Central Africa
11) South America
12) South America
13) India

14) South America

Host species

01), 02), 03), 04), 05), 06), 07), 08) and 09) Certain species of the genus Tetramorium Mayr, 1855

01) T. alpestre Steiner, Schlick-Steiner & Seifert, 2010 and T. impurum (Förster, 1850)
and maybe T. caespitum (Linnaeus, 1758)?
02) T. cf. caespitum (Linnaeus, 1758)
03) T. cf. chefketi Forel, 1911
04) T. cf. chefketi Forel, 1911

05) A species from the genus Tetramorium Mayr, 1855…

06) T. impurum (Förster, 1850), T. caespitum (Linnaeus, 1758), T. immigrans Santschi, 1927, T. staerckei Kratochvíl, in Kratochvíl, Novák, Šnoflák, 1944
and T. moravicum [Kratochvil, in] Novák & Sadil, 1941, T. diomedeum Emery, 1908, T. chefketi Forel, 1911
07) T. immigrans Santschi, 1927

08) T. sericeiventre Emery, 1877 and T. sepositum Santschi, 1918
09) T. avium Bolton, 1980

10), 11), 12), 13) and 14) Certain species of the genus Pheidole Westwood, 1839

10) P. megacephala (Fabricius, 1793) subsp. melancholica Santschi, 1912
11) P. strobeli Emery, 1906
12) P. nitidula Emery, 1888
13) P. indica Mayr, 1879

14) P. obscurior Forel, 1886

A remark about synonymy

Anergates friedlandi Creighton, 1934 is now a synonym from Anergates atratulus (Schenck, 1852), more precisely an introduced form in North Americe. So, the name is Anergates atratulus (Schenck, 1852) or, if you follow Ward, Brady, Fisher, Schultz, 2015 ("2014"), Tetramorium atratulum (Schenck, 1852)...

Synonyms of host species

- T. caespitum (Linnaeus, 1758) (= Formica caespitum Linnaeus, 1758)
- T. impurum (Förster, 1850) (= Myrmica impura Förster, 1850)
- T. staerckei Kratochvíl, in Kratochvíl, Novák, Šnoflák, 1944 (= T. caespitum (Linnaeus, 1758) subsp. hungarica Röszler, 1935 ("1933-34") var. staerckei Röszler, 1936)
- P. megacephala (Fabricius, 1793) (= Formica megecephala Fabricius, 1793)
- P. megacephala (Fabricius, 1793) subsp. melancholica Santschi, 1912 was originally described as P. punctulata Mayr, 1866 st. melancholica Santschi, 1912
- The host species P. strobeli Emery, 1906 was known, during much of its history, as a subspecies of an other species, nl. P. nitidula Emery, 1888 subsp. richteri Forel, 1909 (a much used junior synonym of the older P. perversa Forel, 1908 subsp. richteri Forel, 1909) and known, in 1922, for a while as P. strobeli Emery, 1906 subsp. richteri Forel, 1909, this since Santschi, 1916. The species P. strobeli Emery, 1906 was revived from synonymy (with P. nitidula Emery, 1888 since Wilson, 2003) and the subspecies was synonymized with it in Casadei-Ferreira, Economo, Feitosa, 2020. Or: P. strobeli Emery, 1906 (= P. perversa Forel, 1908 subsp. richteri Forel, 1909)
- P. obscurior Forel, 1886 was revived from synonymy with the 1 page older P. susannae Forel, 1886 and this in Casadei-Ferreira, Economo, Feitosa, 2020.
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Mittwoch 29. Dezember 2021, 19:03

First the list of Teleutomyrmex Literature:

I did put an oldie in it, that is, an oldie for me. I started reading it in 1983 but it was to complicated for me. So I got to "The insect societies" and up to the ants. And so it stayed since then. That is, until now. Read yesterday a part of it for the first time. All the pages with Teleutomyrmex in it. 4 pages. With only one mistake, later changed by more observations.

Wilson, E. O., 1975, “Sociobiology. The new synthesis.” Cambridge, Mass., Harvard University Press, 697 pp.
- 697 pp., ?? figs. - [1975-01-01].
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Mittwoch 29. Dezember 2021, 19:08

An official literature list of Teleutomyrmex Kutter, 1950.

Baroni Urbani, C., 1967, “Le distribuzioni geografiche discontinue dei Formicidi mirmecobiotici.” Archivio Botanico e Biogeografico Italiano, vol. 43, [=(4)12], p. 355-365.

Baur, A., Buschinger, A., Zimmermann, F. K., 1993, “Molecular cloning and sequencing of 18S rDNA gene fragments from six different ant species.” Insectes Sociaux, vol. 40, p. 325-335.

Baur, A., Chalwatzis, N., Buschinger, A., Zimmermann, F. K., 1995, “Mitochondrial DNA sequences reveal close relationships between social parasitic ants and their host species.” Current Genetics, vol. 28, p. 242-247.

Baur, A., Sanetra, M., Chalwatzis, N., Buschinger, A., Zimmermann, F. K., 1996, “Sequence comparisons of the internal transcribed spacer region of ribosomal genes support close relationships between parasitic ants and their respective host species (Hymenoptera: Formicidae).” Insectes Sociaux, vol. 43, p. 53-67.

Bernard, F., 1967 ["1968"], “Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale.” Paris, Masson, 411 pp.

Blaimer, B. B., Ward, P. S., Schultz, T. R., Fisher, B. L., Brady, S. G., 2018, “Paleotropical Diversification Dominates the Evolution of the Hyperdiverse Ant Tribe Crematogastrini (Hymenoptera: Formicidae).” Insect Systematics and Diversity, vol. 2, no. 5, article no. 3, p. 1-14 (+ 2 supporting files.).

Bolton, B., 1976, “The ant tribe Tetramoriini (Hymenoptera: Formicidae). Constituent genera, review of smaller genera and revision of Triglyphothrix Forel.” Bulletin of the British Museum (Natural History) (Entomology), vol. 34, p. 281-379.

Bolton, B., 1994, “Identification guide to the ant genera of the world.” Cambridge, Mass., Harvard University Press, 222 pp.

Bolton, B., 1995, “A taxonomic and zoogeographical census of the extant ant taxa (Hymenoptera: Formicidae).” Journal of Natural History, vol. 29, p. 1037-1056.

Bolton, B., 1995, “A new general catalogue of the ants of the world.” Cambridge, Mass., Harvard University Press, 504 pp.

Bolton, B., 2003, “Synopsis and Classification of Formicidae.” Memoirs of the American Entomological Institute, vol. 71, 370 pp.

Borowiec, L., 2014, “Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae).” Genus (Wroclaw), vol. 25, no. 1-2, p. 1-340.

Bourke, A. F. G., Franks, N. R., 1991, “Alternative adaptations, sympatric speciation and the evolution of parasitic, inquiline ants.” Biological Journal of the Linnean Society, vol. 43, no. 3, p. 157-178.

Brown, W. L., Jr., 1973, “A comparison of the Hylean and Congo-West African rain forest ant faunas.” P. 161-185 in: Meggers, B. J., Ayensu, E. S., Duckworth, W. D. (editors.), 1973, “Tropical forest ecosystems in Africa and South America: a comparative review.” Smithsonian Institution Press, Washington, DC., viii + 350 pp.

Brun, R., 1952, “Das zentralnervensystem von Teleutomyrmex schneideri Kutt. (Hym. Formicid.). III. Mitteilung.” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 25, no. 2, p. 73-86.

Brun, R., 1959, “Le cerveau des fourmis et des insectes en général comme instrument de formation des réflexes conditionnés.” P. 11-25 in: International Union of Biological Sciences, Union Internationale des Sciences Biologiques, Experimental Psychology and Animal Behaviour Section, 1959, “Animal psychology seminars: Strasbourg University, October 1956, and Brussels University, August 1957.” London, Pergamon Press, 148 pp.

Buschinger, A., 1970, “Neue Vorstellungen zur Evolution des Sozialparasitismus und der Dulosis bei Ameisen (Hym., Formicidae).” Biologisches Zentralblatt, vol. 89, no. 3, p. 273-299.

Buschinger, A., 1971, “Zur Verbreitung und Lebensweise sozialparasitischer Ameisen des Schweizer Wallis (Hym., Formicidae).” Zoologischer Anzeiger, vol. 186, no. 1/2, p. 47-59.

Buschinger, A., 1974, “Monogynie und Polygynie in Insektensozietäten.” P. 862-896 in: Schmidt, G. H. (editor.), 1974, “Sozialpolymorphismus bei Insekten. Probleme der Kastenbildung im Tierreich.” Stuttgart, Wissenschaftliche Verlagsgesellschaft mbH, xxiv + 974 pp.

Buschinger, A., 1974, “Polymorphismus und Polyethismus sozialparasitischer Hymenopteren.” P. 897-934 in: Schmidt, G. H. (editor.), 1974, “Sozialpolymorphismus bei Insekten. Probleme der Kastenbildung im Tierreich.” Stuttgart, Wissenschaftliche Verlagsgesellschaft mbH, xxiv + 974 pp.

Buschinger, A., 1985, “New records of rare parasitic ants (Hym., Form.) in the French Alps.” Insectes Sociaux, vol. 32, no. 3, p. 321-324.

Buschinger, A., 1986, “Evolution of social parasitism in ants.” Trends in Ecology and Evolution, vol. 1, no. 6, p. 155-160.

Buschinger, A., 1987, “Teleutomyrmex schneideri Kutter, 1950 and other parasitic ants found in the Pyrenees (Hymenoptera, Formicidae).” Spixiana, vol. 10, no. 1, p. 81-83.

Buschinger, A., 1990, “Evolutionary transitions between types of social parasitism in ants, hypotheses and evidence.” P. 145-146 in: Veeresh, G. K., Mallik, B., Viraktamath, C. A. (editors.), 1990, “Social insects and the environment. Proceedings of the 11th International Congress of IUSSI, 1990.” New Delhi, Oxford & IBH Publishing Co., xxxi + 765 pp.

Buschinger, A., 1990, “Sympatric speciation and radiative evolution of socially parasitic ants. - Heretic hypotheses and their factual background.” Zeitschrift für Zoologische Systematik und Evolutionsforschung, vol. 28, p. 241-260.

Buschinger, A., 1995, “Nicht am Ende: Die “Endameise” Teleutomyrmex schneideri.” Ameisenschutz aktuell, vol. 9, p. 1-7.

Buschinger, A., 1999, “Wiederfund der sozialparasitischen Ameise Teleutomyrmex schneideri in der Schweiz.” Mitteilungen der Schweizerischen Entomologischen Gesellschaft, vol. 72, p. 277-279.

Buschinger, A., 2000, “Die “Endameise” Teleutomyrmex schneideri in der Schweiz: Erster Wiederfund nach 50 Jahren.” Ameisenschutz aktuell, vol. 14, p. 43-45.

Buschinger, A., 2009, “Social parasitism among ants: a review (Hymenoptera: Formicidae).” Myrmecological News, vol. 12, p. 219-235 (+ 1 supporting file.).

Casevitz-Weulersse, J., Galkowski, C., 2009, “Liste actualisée des fourmis de France (Hymenoptera, Formicidae).” Bulletin de la Société Entomologique de France, vol. 114, no. 4, p. 475-510.

Collingwood, C. A., 1956, “Ant hunting in France.” Entomologist, vol. 89, p. 105-108.

Collingwood, C. A., 1956, “A rare parasitic ant (Hym., Formicidae) in France.” Entomologist's Monthly Magazine, vol. 92, p. 197.

Csősz, S., Radchenko, A., Schulz, A., 2007, “Taxonomic revision of the Palaearctic Tetramorium chefketi species complex (Hymenoptera: Formicidae).” Zootaxa, vol. 1405, p. 1-38.

Cuesta, D., García, F., García-Tejero, S., Espadaler, X., 2009, “Resumen charla: Aportaciones a la biología de Teleutomyrmex schneideri Kutter, 1950: primer caso de cría en cautividad. [Contributions to the biology of Teleutomyrmex schneideri Kutter, 1950: first case of keeping in captivity.]” [abstract]. Iberomyrmex, vol. 1, p. 24.

Cuesta-Segura, A.D., Espadaler, X., Garcia Garcia, F., 2017, “Hormigas de los brezales de Calluna cantábricos (NO España) (Hymenoptera: Formicidae). [Ants of the Cantabrian Calluna-heathlands (NW Spain) (Hymenoptera: Formicidae).]” Iberomyrmex, vol. 9, p. 25-43.

Cuesta-Segura, A. D., García García, F., Catarineu, C., García-Tejero, S., Espadaler, X., 2018, “Actualización de la Distribución y Hospedadores de la Hormiga Parasita Teleutomyrmex schneideri Kutter, 1950 en la Peninsula Ibérica (Hymenoptera: Formicidae).” Boletín de la Sociedad Entomológica Aragonesa (S.E.A.), nº. 63 (31/12/2018), p. 235–239.

de la Mora, A., Sankovitz, M., Purcell, J., 2020, "Ants (Hymenoptera: Formicidae) as host and intruder: recent advances and future directions in the study of exploitative strategies." Myrmecological News, vol. 30, p. 53-71 (+ 1 supporting file.).

Dlussky, G. M., Fedoseeva, E. B., 1988 (“1987”), “[The origin and early stages of evolution of ants (Hymenoptera: Formicidae).]” P. 70-144 in: Ponomarenko, A. G. (editor.), “[The Creaceous biocenotic crisis and the evolution of insects.]” [In Russian.]. Nauka, Moscow. 231 pp.

Dlussky, G. M., Soyunov, O. S., Zabelin, S. I., 1990 [“1989”], “Muravji Turkmenistana.”, or “[Ants of Turkmenistan.]” [In Russian.]. Ashkabad, Ylym Press, 275 pp.

Dobrzanska, J., Dobrzanski, J., 1988, “Pasozytnictwo spoleczne u mrowek. Pasozytnictwo obowiazkowe.”, or “[Social parasitism in ants. Obligatory parasitism.]” Kosmos, vol. 37, no. 4, p. 617-639.
-> Dobrzanska, J., Dobrzanski, J., 1988, “1. Inkwilinizm.”, or “[Inquilinism.]” Kosmos, vol. 37, no. 4, p. 617-629.
-> Dobrzanska, J., Dobrzanski, J., 1988, “2. Niewolnictwo.”, or “[Slavery.]” Kosmos, vol. 37, no. 4, p. 629-639.

Dumpert, K., 1978, “Das Sozialleben der Ameisen.” (First Edition.). Pareys Studientexte, nr. 18. Mit 95 Abbildungen. Verlag Paul Parey, Berlin und Hamburg, 1977, 253 pp.

Dumpert, K., 1994, “Das Sozialleben der Ameisen. 2., neubearbeitete Auflage.” (Second Edition.). Pareys Studientexte, nr. 18. Mit 94 Abbildungen. Verlag Paul Parey, Berlin und Hamburg, 1994, 257 pp.

Espadaler, X., Cuesta, D., 2006, “Notas / Notes. Teleutomyrmex schneideri Kutter, 1950 en España (Hymenoptera, Formicidae).” Graellsia, vol. 62, no. 2, p. 261-262.

Espadaler, X., Gómez, K., 2014, “Tetramorium biskrense Forel, 1904 en España y Portugal peninsulares (Hymenoptera, Formicidae).” Boletín de la Sociedad entomológica Aragonesa (SEA), vol. 55, p. 303-305.

García, F., 2019, “Parásitos y myrmecófilos. Hormigas parásitas sociales ibéricas.” LaMarabunta digital, Nueva Edición, Número 4, Junio 2019, p. 27-44.

Gösswald, K., 1952, “Zur Biologie und Histologie parasitär degenerierter Ameisenarten mit besonderer Berücksichtigung von Teleutomyrmex schneideri Kutter (Tribus Tetramorini).” P. 446-448 in: de Wilde, J. (editor.), 1952, “Transactions of the IX-th International Congress of Entomology, Amsterdam, August 17-24 (1951), vol. 1.” The Netherlands, The Hague, Exselsior’s Photo-Offset, lxi + 1116 pp.
- His name is spelled “Göszwald” in this publication!

Gösswald, K., 1953, “Histologische Untersuchungen an der arbeiterlosen Ameise Teleutomyrmex schneideri Kutter (Hym. Formicidae).” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 26, no. 2, p. 81-128.

Gösswald, K., 1954, “Unsere Ameisen. I. Teil.” Kosmos-Bändchen I, Beilage 204. Stuttgart, Franckh'sche Verlagshandlung, 88 pp.

Gösswald, K., 1955, “Unsere Ameisen. II. Teil.” Kosmos-Bändchen II, Beilage 206. Stuttgart, Franckh'sche Verlagshandlung, 80 pp.

Gösswald, K., 1985, “Organisation und Leben der Ameisen.” Stuttgart, Wissenschaftliche Verlagsgesellschaft mbH, 355 pp.

Hölldobler, B. K., Wilson, E. O., 1990, “The Ants.” Cambridge, Mass., Belknap Press of Harvard University Press, xiv + 732 pp.

Hölldobler, B. K., Wilson, E. O., 1994, “Journey to the Ants: A Story of Scientific Exploration.” Cambridge, Mass., Belknap Press of Harvard University Press, 304 pp.

Kiran, K., Karaman, C., Lapeva-Gjonova, A., Aksoy, V., 2017, “Two new species of the “ultimate“ parasitic ant genus Teleutomyrmex Kutter, 1950 (Hymenoptera: Formicidae) from the Western Palaearctic.“ Myrmecological News, vol. 25, p. 145-155.

Kusnezov, N., 1954, “Phyletische Bedeutung der Maxillar- und Labialtaster der Ameisen.” Zoologischer Anzeiger, vol. 153, no. 1/2, p. 28-38.

Kutter, H., 1950, “Über eine neue, extrem parasitische Ameise. 1. Mitteilung.” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 23, no. 2, p. 81-94.

Kutter, H., 1950, “Über zwei neue Ameisen.” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 23, no. 3, p. 337-346.

Kutter, H., 1950, “Über Doronomyrmex und verwandte Ameisen. 2. Mitteilung.” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 23, no. 3, p. 347-353.

Kutter, H., 1951, “Von dufttäuschenden Mörderinnen und berittener Königin.” Du: kulturelle Monatsschrift, vol. 11, no. 4, p. 44-47.

Kutter, H., 1952, “Über Plagiolepis xene Stärcke (Hym. Formicid.).” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 25, no. 2, p. 57-72.

Kutter, H., 1963, “Miscellanea myrmecologica I.” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 36, no. 1 and 2, p. 129-137.

Kutter, H., 1968 ("1967"), “Liste sozialparasitischer Ameisen.” Archives. Institut Grand-Ducal de Luxembourg (n.s.), vol. 33, p. 201-210.

Kutter, H., 1968 (“1969”), “Die sozialparasitischen Ameisen der Schweiz.” Neujahrsblatt herausgegeben von der Naturforschenden Gesellschaft in Zürich auf das Jahr 1969, vol. 171, (Ausgegeben am 31. Dezember 1968), p. 1-62.

Kutter, H., 1977, “Hymenoptera, Formicidae.” Insecta Helvetica. Fauna, vol. 6, p. 1-298.

Kutter, H., 1978, “Hymenoptera, Formicidae.” Insecta Helvetica. Fauna, vol. 6a, unpaginated, illustr.

Meyer, G., 1951, “Histologische Untersuchungen an einer arbeiterlosen Ameise Anergates atratulus Schenck.” Inaugural-Dissertation, Institut für Angewandte Zoologie, Würzburg.

Meyer, G. F., 1955, “Untersuchungen an einer parasitischen Ameise (Anergates atratulus Schenck).” Insectes Sociaux, vol. 2, no. 2, p. 163-171.

Passera, L., 1984, “L'Organisation Sociale des Fourmis.” Toulouse, Editions Privat, 360 pp.

Rabeling, C., 2020, “Social Parasitism.” P. 836-858 in: Starr, C. K. (editor.), 2021, “Encyclopedia of Social Insects.” Springer International Publishing, Cham., xxvi + 1049 pp.

Reyes López, J., Benavente Martínez, A., 2011, “Notas Científicas: Nueva cita de Teleutomyrmex kutteri Tinaut, 1990 (Hym., Formicidae) para la Península Ibérica. New record of Teleutomyrmex kutteri Tinaut, 1990 (Hym., Formicidae) from the Iberian Peninsula.” Boletín de la Sociedad Entomológica Aragonesa (S.E.A.), nº. 49, p. 206.

Sanetra, M., 1996, “Systematik, Biologie und Phylogenie westpaläarktischer Ameisen der Tribus Tetramoriini unter besonderer Berücksichtigung molekularer Merkmale (Hymenoptera: Formicidae).” Thesis, Technische Hochschule Darmstadt, p. 1-229.

Sanetra, M., Buschinger, A., 2000, “Phylogenetic relationships among social parasites and their hosts in the ant tribe Tetramoriini (Hymenoptera: Formicidae).” European Journal of Entomology, vol. 97, no. 1, p. 95-117.

Sanetra, M., Felger, S., Buschinger, A., Zimmermann, F. K., 1998, “On the evolutionary history of social parasites in the ant tribe Tetramoriini (Hymenoptera: Formicidae).” P. 413 in: Schwarz, M. P., Hogendoorn, K. (editors.), 1998, “Social insects at the turn of the millennium. Proceedings of the XIII International Congress of IUSSI. Adelaide, Australia. 29 December 1998 - 3 January 1999.” Adelaide, XIII Congress of IUSSI, 535 pp.

Sanetra, M., Heinze, J., Buschinger, A., 1994, “Enzyme polymorphism in the ant genus Tetramorium Mayr and its social parasites (Hymenoptera: Formicidae).” Biochemical Systematics and Ecology, vol. 22, p. 753-759.

Schlick-Steiner, B. C., Steiner, F. M., Moder, K., Seifert, B., Sanetra, M., Dyreson, E., Stauffer, C., Christian, E., 2006, “A multidisciplinary approach reveals cryptic diversity in Western Palaearctic Tetramorium ants (Hymenoptera: Formicidae).” Molecular Phylogenetics and Evolution, vol. 40, p. 259-273.

Schönitzer, K., Lawitzky, G., 1987, “A phylogenetic study of the antenna cleaner in Formicidae, Mutillidae, and Tiphiidae (Insecta, Hymenoptera).” Zoomorphology (Berlin), vol 107, no. 5, p. 273-285.

Seifert, B., 1996, “Ameisen: beobachten, bestimmen.” Naturbuch Verlag, Augsburg, 351 pp.

Seifert, B., 2007, “Die Ameisen Mittel- und Nordeuropas.” Lutra Verlags- und Vertriebsgesellschaft, Tauer/Klitten, 368 pp.

Seifert, B., 2018, “The ants of Central and North Europe.” Lutra Verlags- und Vertriebsgesellschaft, Tauer/Boxberg, 407 pp.

Seifert, B., Buschinger, A., Aldawood, A., Antonova, V., Bharti, H., Borowiec, L., Dekoninck, W., Dubovikoff, D., Espadaler, X., Flegr, J. , Georgiadis, C., Heinze, J., Neumeyer, R., Ødegaard, F., Oettler, J., Radchenko, A., Schultz, R., Sharaf, M., Trager, J., Vesnić, A., Wiezik, M., Zettel, H., 2016, “Banning paraphylies and executing Linnaean taxonomy is discordant and reduces the evolutionary and semantic information content of biological nomenclature.” Insectes Sociaux, vol. 63, no. 2, p. 237-242.

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Wegnez, P., Ignace, D., Lommelen, E., Hardy, M., Bogaert, J., Nilsson, C., 2015, “Redécouverte de Teleutomyrmex schneideri Kutter, 1950 dans les Alpes françaises (Hymenoptera: Formicidae).” Bulletin de la Société royale belge d’Entomologie/Bulletin van de Koninklijke Belgische Vereniging voor Entomologie, vol. 151, p. 52-57.

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104 references.
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Mittwoch 29. Dezember 2021, 19:16

And in the list "not in" Teleutomyrmex and Anergates:

I added 4 publications, 1 from Belgium, 1 from Deutschland, 1 from England and 1 from U.S.A.

Billen, J. P. J. (editor.), 1992, “Biology and evolution of social insects.” Leuven, Leuven University Press, ix + 390 pp.

Eder, J., Rembold, H. (editors.), 1987, “Chemistry and biology of social insects.” München, Verlag J. Peperny, xxxv + 757 pp.

Howse, P. E., Clement, J.-L. (editors.), 1981, “Biosystematics of social insects.” Systematics Association Special Volume No. 19. London, Academic Press, 346 pp.

Lofgren, C. S., Vander Meer, R. K. (editors.), 1986, “Fire ants and leaf-cutting ants. Biology and management.” Boulder, Westview Press, xv + 435 pp.

P.S. I have all of them except Forel 1921-1948!
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